
Aluminum toxicity usually
damages the root system first,
while the tops may look normal or
may appear drought stress, P or
Ca deficiency. Aluminum-affected
roots tend to be shortened and
swollen, having a stubby
appearance (Figure 9). A high level
of Al impairs root elongation and
decreases nutrient uptake; it
interferes with cell division at the
root apex, increases the rigidity of
the cell wall by crosslinking of
pectins which usually carry
negative charge, and reduces DNA
replication because of increased
rigidity of the double helix (Gupta
et al. 2013; Eekhout et al. 2017;
Bojorquez-Quintal et al. 2017).
Figure 9. Aluminum effect on roots. Sesbania seedlings grown in an
Ultisol (non-amended pH 4.2, right; and limed pH 5.5, left) of Hawaii.
3.2 Manganese Toxicity
Some soils in the tropics, particularly those of the Oxisol order, can contain high levels of Mn. For
example, the Wahiawa series, Oxisol order, in Hawaii has 1.2 – 1.6% total Mn mostly as MnO
2
(Hue et al.
2001). For comparison, background levels of total Mn in world’s soils average about 0.05% (500 mg/kg dry
weight) (WHO 2004). Under acidic conditions and with the supply of electron (e-) from SOM, MnO
2
will
dissolve into soluble Mn
2+
according to the reaction:
MnO
2
+ 4H
+
+ 2e
-
↔ Mn
2+
+ 2H
2
O R.23
Equilibrium constant of R.23 can be expressed as:
K = (Mn
2+
)/{(H
+
)
4
*(e
-
)
2
} R.24
If we assume that the system is poised, meaning log (H
+
) + log(e
-
) constant, which is often the case in
soils (Lindsay 1979), then R.24 becomes
Log(Mn
2+
) = constant -2pH (Hue and Mai 2002) R.25
R.25 would predict that for every pH unit decrease, (Mn
2+
) activity (and concentration) would
increase by 100 fold. In reality, however, because soil solution may contain other inorganic and organic
ions/molecules that can complex Mn
2+
and keep more Mn
2+
in solution regardless of pH, Mn
2+
only
increases about 10 fold for each pH unit drop as shown in Figure 10.