College of Tropical Agriculture and Human Resources
Dr. Arnold H. Hara




  • Hara, A.H., R.Y. Niino-DuPonte, S.K. Cabral, K.L. Aoki, and J. Zarders. 2013. Best Management Practices: Coqui frog Eleutherodactylus coqui. University of Hawaii at Manoa, CTAHR, Cooperative Extension Service, Hilo, HI.
  • Marr, S.R., S.A. Johnson, A.H. Hara, and M.E. McGarrity. 2010. Preliminary evaluation of the potential of the helminth parasite Rhabdias elegans as a biological control agent for invasive Puerto Rican coquis, Eleutherodactylus coqui, in Hawaii. Biological Control 54: 69-74.
  • Hara, A.H., C.M. Jacobsen, S.R. Marr and R.Y. Niino-DuPonte. 2010. Hot water as a potential disinfestation treatment for an invasive anuran amphibian, Eleutherodactylus coqui Thomas (Leptodactylidae) on potted plants. International Journal of Pest Management 56(3): 255-263.
  • Hara, A.H. and R.Y. Niino-DuPonte. 2010c. UH coqui frog research details control methods. Hawaii Landscape, Jan/Feb 2010, Landscape Industry Council of Hawai`i.
  • Hara, A.H. and R.Y. Niino-DuPonte. 2010b. Tips for outwitting the wily coqui frog. Hawaii Tribune Herald, Jan. 10, 2010. Hilo, HI.
  • Hara,A.H. and R.Y. Niino-DuPonte. 2010a. There are several ways to control the coqui frog. Hawaii Tribune Herald, Jan. 3, 2010. Hilo, HI.
  • Marr, S.R., W.J. Mautz and A.H. Hara. 2008. Parasite loss and introduced species: a comparison of the parasites of the Puerto Rican tree frog, Eleutherodactylus coqui, in its native and introduced ranges. Bioloigcal Invasions 10: 1289-1298.
  • Woolbright, L., A.H. Hara, C.M. Jacobsen, W.J. Mautz, and F.B. Benevides, Jr. 2006.  Population densities of the coqui frog Eleutherodactylus coqui (Anura:Leptodactylidae) in newly invaded Hawai`i and in native Puerto Rico.  J. of Herpetology 40(1): 122-128.
  • Hara, A.H. 2004.  "Coqui Frog Invasion in Hawai`i" (VIDEO)
    • Video Evaluation Form (Fillable)
  • Chun, S., A.H. Hara, and R.Y. Niino-DuPonte. 2003. Greenhouse frog or a coqui frog? Differences between two Eleutherdactylus species found in Hawai`i. Cooperative Extension Service. University of Hawai`i CTAHR OSC. Miscellaneous Pests MP-3. 1 oversized page.
  • Coqui Frog Working Group.  2003. Coqui frog control for homeowners. Cooperative Extension Service, University of Hawai`i CTAHR OSC. Miscellaneous Pests MP-5. 4-fold, 2 pp. (rev 11/04, rev 5/06).
  • Hara, A.H. 2003.  "Control of Coqui Frogs in Hawai`i" (Website) http://www.ctahr.hawaii.edu/coqui/


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